
姓名:刘煜
职称:研究员
职务:云南省古生物研究重点实验室主任
邮箱: yu.liu@ynu.edu.cn;zac.liu@126.com
地址:云南省昆明市五华区翠湖北路2号云南大学东陆校区科学馆305A
邮编:650500
https://www.researchgate.net/profile/Yu_Liu6
http://www.yklp.ynu.edu.cn/info/1063/1306.htm
本团队常年招募博士后,有意者请发送邮件至:yu.liu@ynu.edu.cn
个人简介
刘煜,男,汉族,1980年生。理学博士,云南大学古生物研究院云南省古生物研究重点实验室研究员,云南大学海外引进人才。从事节肢动物演化的相关研究。
自2006年在欧盟玛丽居里基金会提供的全额奖学金的资助下远赴德国沃尔姆大学(Universität Ulm)攻读博士学位开始,刘煜在德国自然科学界从事早期生命科学研究和教育工作逾10年。他于2016年上半年通过了德国教育系统最高级别的教学及科研考核(Habilitation),获得Privatdozent头衔。这一头衔同时也赋予其在德国境内任一高校申请全职教授的资格。刘煜于2016年10月作为云南大学海外引进人才正式进入云南省古生物研究重点实验室工作并于次年5月入选国家海外高层次青年人才项目。
过去10年间,刘煜的研究方向涵盖了胚胎发育学、神经发育生物学及古生物学等研究动物进化事件的国际前沿领域,并在各领域均有若干SCI学术论文发表。截至目前,共发表SCI英文文章60余篇,英文书本章节1篇。其中,第一作者或(共同)通讯作者文章20余篇。代表性论文分别发表于PNAS,Nature Communications,Current Biology,Proceedings of the Royal Society B: Biological Sciences等中科院JCR一区期刊或Nature Index期刊上。主要工作内容为使用先进成像方法深入研究早寒武世澄江动物群及其它保存类型的节肢动物化石。
近年来,刘煜主要致力于将目前国际生物学研究领域中最前沿的多种先进研究手段和方法创新性地引入到澄江动物群化石标本的研究当中。经其研究工作创新性地引入的先进技术手段主要有:荧光显微镜、电子扫描显微镜和显微CT。前二者主要用于提取普通光学显微镜下无法观察到的保存在化石表面的动物身体结构信息;后者则主要用于提取保存在化石内部的动物身体三维结构信息。刘煜于2015年在Scientific Reports上发表文章将显微CT技术首次引入到澄江生物群化石研究中。显微CT扫描是无损研究技术,不仅可以让研究者观察到被围岩包埋或被动物化石的其它身体结构覆盖的、用钢针手工处理难以暴露的精细构造,而且还极大程度地保护了作为世界自然遗产的珍贵寒武纪澄江化石的完整性,使这些化石在研究过程中不受到人为(如:钢针修理)的破坏。刘煜的工作为澄江节肢动物化石的研究打开了多个方面的窗口(如:古发育生物学、古神经解剖学及古生态学等)。在借助上述技术手段详细了解了寒武纪早期各种动物的形态细节的基础上,古生物学家和进化生物学家们就可以对这些分支学科进行深入研究,最终实现对‘寒武纪大爆发’这一曾困惑了达尔文多年的早期动物进化事件形成更深入和全面的认识。

早寒武世澄江生物群部分节肢动物化石及三维模型
迷人林乔利虫(Leanchoilia illecebrosa)7毫米幼虫三维模型
罗氏小虾(Xiaocaris luoi)腹部腿肢细节三维模型
多节佴材村虫(Ercaicunia multinodosa)三维模型
学习和工作经历
1998.09–2002.07 云南大学,理学学士(固体地球物理学)
2002.09–2005.07 云南大学,理学硕士(古生物学与地层学)
2006.04–2009.07 德国沃尔姆大学,博士(自然科学)
2009.12–2016.03 德国慕尼黑大学,副教授(发育神经生物学)
2016.04–2016.09 德国慕尼黑大学,副教授(古生物学与地层学)
2016.10–今 云南大学古生物研究院,云南省古生物研究重点实验室
2019.10–2020.10 哈佛大学动物与演化生物学系,访问学者
行政职务
2017.05–2022.05 云南大学云南省古生物研究重点实验室,常务副主任
2020.01–2025.04 云南大学古生物研究院,副院长
2022.06起至今 云南省古生物研究重点实验室,主任
承担项目情况
1. 云南省委组织部,2023年度“云岭学者”专项竞争性项目,2024.01.01-2026.12.31,250万元,在研,主持
2. 云南省科技厅,云南省基础研究专项重大项目,202401BC070012,三维视野下的“寒武纪生命大爆发”研究,2024.03.01-2027.02.28,300万元,在研,主持
3. 云南省科技厅,云南省科技厅基础研究重点项目,202301AS070049,澄江动物群节肢动物发育生物学研究,2023.06-2026.05,50万,在研,主持
4. 国家自然科学基金委,国际(地区)合作与交流项目,41861134032,澄江生物群节肢动物的精细古生物学研究,2019. 01至2021. 12,133万元,在研,参加
5. 云南省科技厅,应用基础研究计划重点项目,2018FA025,寒武纪大爆发时期的节肢动物生态学研究,2018. 06至2021. 05,50万元,已结题,主持
6. 云南省科技厅,国际科技合作项目,2018IA073,云南省古生物化石国际联合研究中心,2018. 01至2019. 12,50万元,已结题,主持
7. 中组部,中组部国家海外高层次青年人才项目,2017. 07至2020. 06,300万元,已结题,主持
8. 国家自然科学基金委,海外及港澳台学者合作研究项目,41528202,澄江动物群节肢动物古发育生物学及古生态学研究,2016. 01至2017. 12,20万元,已结题,主持
9. 德国慕尼黑大学,优秀青年学者基金项目,早寒武世澄江动物群中节肢动物生态学、发育生物学及神经生物学研究,2015. 02至2016. 09,2.5万欧元,已结题,主持

云南大学寒武纪节肢动物研究团队合影(摄于2019年10月)

云南大学节肢动物演化研究团队部分成员合影(摄于2021年9月)
奖励情况
2020年,国家自然科学奖二等奖(排名第五)
2017年,云南省“云岭高层次人才”(2019年更名为“云南省千人计划”)
2016年,发表于PNAS的科研成果入选“中国古生物学十大进展”
2011年,获云南省科学技术厅颁发“云南省科学技术奖”特等奖(排名第三)
论文著作
1. Peng, A., Engel, M. S., Boderau, M., Legendre, F., Liu, Y., Nyunt, T. T., Wang, B.*, Nel, A*. 2025. Descending from trees: a Cretaceous winged ice-crawler illuminates the ecological shift and origin of Grylloblattidae. Proceedings of the Royal Society B: Biological Sciences. 292, 20250557.https://doi.org/10.1098/rspb.2025.0557
2. Zhuang, Y., Luo, C., Tang, D., Araújo, J. P. M. de, Yu, H., Zhao, J., Haelewaters, D., Nyunt, T. T., Zhang, Q., Jarzembowski, E. A., Meng, G., Ran, H., Liu, Y., Wang, B*. 2025. Cretaceous entomopathogenic fungi illuminate the early evolution of insect–fungal associations. Proceedings of the Royal Society B: Biological Sciences. 292, 20250407. http://doi.org/10.1098/rspb.2025.0407
3. Lustri, L.*, Collantes, L., Esteves, C. J. P., O’Flynn, R. J., Saleh, F., Liu, Y. 2025. A Hypothesis on Suspension Feeding in Early Chelicerates (Offacolidae). Diversity. 17 (6), 412. https://doi.org/10.3390/d17060412
4. Zhuang, Y., Jouault, C., Perrichot, V., Wang, B., Liu, Y.*, Zhang, Q.*. 2025. Unveiling the diversity of iron maiden ants: morphological insights from two new Cretaceous species. Zoological Journal of the Linnean Society. 204 (2), zlaf044. https://doi.org/10.1093/zoolinnean/zlaf044
5. Wu, Z., Dunlop, J.A., Klimov, P.B., Mai, H., Peng, A., Liu, Y.*. 2025. A new whip scorpion (Arachnida: Thelyphonida) with a phoretic mite (Acariformes: Trochometridiidae) from Mid-Cretaceous Kachin amber. BMC Ecol Evo. 25, 55. https://doi.org/10.1186/s12862-025-02392-w
6. Zhuang, Y., Perrichot, V., Li, J., Zhuo, D., Liu, Y.*, Wang, B., Zhang, Q.*, Boudinot, B.E. 2025. Systematics and early evolution of ants (Hymenoptera: Formicidae): the first male and worker-queen association for †Gerontoformica. Myrmecological News. 35, 89-98. DOI: 10.25849/myrmecol.news_035:089
7. Schmidt, M., Schoenemann, B.*, Hou, X., Melzer, R.R., Liu, Y*. 2025. Pygmaclypeatus daziensis, a unique lower Cambrian arthropod with two different compound eye systems.Communications Biology. 8, 317. https://doi.org/10.1038/s42003-025-07664-1
8. Schmidt, M., Melzer, R. R., Mai, H., Hou, X., Liu, Y*. 2025. Drishti and Amira – different visualizations exemplified by the early Cambrian Chengjiang arthropod Leanchoilia illecebrosa from China. Palaeontologia Electronica, 28(1):a5. https://doi.org/10.26879/1358.
9. O’Flynn, RJ., Williams, M.*, Liu, Y.*, Hou, X., Guo, J., Edgecombe, GD. The early Cambrian Kuamaia lata, an artiopodan euarthropod with a raptorial frontal appendage. Journal of Paleontology. 2024;98(5):808-820. doi:10.1017/jpa.2024.33
10. Parry, L.A.*, Briggs, D.E.G., Ran, R., O’Flynn, R.J., Mai, H., Clark, E.G., Liu, Y*. 2024. A pyritized Ordovician leanchoiliid arthropod. Current Biology. 34,1-9. https://doi.org/10.1016/j.cub.2024.10.013.
11. Zhang M., Liu Y.*, Mai H., Schmidt M., Hou X. 2024. Ventral Morphology and Ecological Implications of Cindarella eucalla (Artiopoda, Xandarellida) from Chengjiang Biota, China. eLife.13:e100729.
http://.https://doi.org/10.7554/eLife.100729.1.
12. Schmidt, M., Hou, X., Mai, H.,Zhou,G.,Melzer, R.R.,Zhang, X.,Liu,Y.*.2024. Unveiling the ventral morphology of a rare early Cambrian great appendage arthropod from the Chengjiang biota of China. BMC Biology. 22(96). https://doi.org/10.1186/s12915-024-01889-y
13. O'Flynn, R. J., Williams, M., Yu, M., Guo, J., Audo, D., Schmidt, M., Mai, H., Liu, Y.*, Edgecombe, G. D*. 2024. The early Cambrian Bushizheia yangi and head segmentation in upper stem-group euarthropods. Papers in Palaeontology, 10: e1556. https://doi.org/10.1002/spp2.1556.
14. Peng, A., Engel, MS, Liu, Y, Nel, A*. A new lineage of pygidicranid earwigs from mid-Cretaceous amber of northern Myanmar (Dermaptera: Pygidicranidae). Cretaceous Research. 2024;158:105849. doi:10.1016/j.cretres.2024.105849
15. O'Flynn, R.J., Liu, Y.*, Hou, X., Mai, H., Yu, M., Zhuang, S., Williams, M.,Guo, J., Edgecombe, G.D.*.2023. The early Cambrian Kylinxia zhangi and evolution of the arthropod head. Current Biology. 33(18), 4006-4013. https://doi.org/10.1016/j.cub.2023.08.022.
16. Zhang, C., Liu, Y.*, Ortega-Hernández, J.*, Wolfe J.M., Jin , C., Mai, H., Hou, X., Guo, J., Zhai, D.*.2023.Three-dimensional morphology of the biramous appendages in Isoxys from the early Cambrian of South China, and its implications for early euarthropod evolution. Proceedings of the Royal Society B: Biological Sciences. 290, 2023335. http://doi.org/10.1098/rspb.2023.0335.
17. Peng,S.*,Babcock,L.E.,Yang,X.*,Zhu,X.,Liu,Y.2023.A new dameselloid trilobite from the Fulu Biota, Longha Formation (Cambrian: Guzhangian), Yunnan, South China, and revised classification of dameselloids. Palaeoworld.http://doi.org//10.1016/j.palwor.2023.01.006.
18. Tang,Y.,Peng,A.,Wu,Z.,Engel,M.S.,Yang, Z.*, Liu, Y.* .2023. Mygalomorph spiders in mid-Cretaceous Kachin amber (Araneae: Mygalomorphae), northern Myanmar: A new genus and species of the family Macrothelidae. Cretaceous Research, 147, 105514. https://doi.org/10.1016/j.cretres.2023.105514.
19. Zhuang,Y.,Xu,W.,Zhang,G.,Mai,H.,Li,X.,He,H.,Ran,H.*,Liu,Y.*.2022.Unparalleled details of soft tissues in a Cretaceous ant. BMC Ecology and Evolution. 22, 146.https://doi.org/10.1186/s12862-022-02099-2.
20. Li, Y., Zhou, Z.*, Zhai, D., O'Flynn, R.J., Tang, J., Xu, J., Zhou, X., Liu, Y.*.2022. Hydrodynamics and sliding posture analysis of the Cambrian arthropod Ercaicunia multinodosa, Palaeoworld, 32 (3), 385–395.https://doi.org/10.1016/j.palwor.2022.09.005.
21. Zhang, X., Liu, Y.*, O'Flynn, R.J., Schmidt, M., Melzer, R.R., Hou, X., Guo, J., Yu, M., Ortega-Hernández, J.*. 2022. Ventral organization of Jianfengia multisegmentalis Hou, and its implications for the head segmentation of megacheirans. Palaeontology, e12624.https://doi.org/10.1111/pala.12624.
22. Peng, A., Engel, M.S., Zhuang, Y., Wu, Z., Feng, C., Liu, Y.* .2022. A new genus of striped earwigs allied to Zigrasolabis in mid-Cretaceous Kachin amber (Dermaptera: Labiduridae). Cretaceous Research, 139, 105305.https://doi.org/10.1016/j.cretres.2022.105305.
23. Zhang, M., Liu, Y.*, Hou, X., Ortega-Hernández, J.*, Mai, H., Schmidt, M., Melzer, R.R., Guo, J. 2022. Ventral Morphology of the Non-Trilobite Artiopod Retifacies abnormalis Hou, Chen & Lu, 1989, from the Early Cambrian Chengjiang Biota, China. Biology, 11(8), 1235. https://doi.org/10.3390/biology11081235.
24. Yang, X., Peng, S., Babcock L.E., Zhu, X., Liu, Y. 2022. New species of Liostracina Monke, 1903 (Trilobita, Cambrian) from Yunnan, China: Complete holaspid exoskeleton and implications for higher level classification. Journal of Paleontology, first view, 1-16. https://doi.org/10.1017/jpa.2022.56.
25. O'Flynn, R.J., Williams, M.,Yu, M., Harvey, T.H.P., Liu, Y.*. 2022. A new euarthropod with large frontal appendages from the early Cambrian Chengjiang biota. Palaeontologia Electronica, 25(1): a6. https://doi.org/10.26879/1167.
26. Schmidt, M.*, Hou, X., Melzer, R.R.*, Zhai, D., Mai, H., Belojević, J, Chen, X., Ortega-Hernández, J.*, Liu, Y.*. 2022. Before trilobite legs Pygmaclypeatus daziensis reconsidered and the ancestral appendicular organization of Cambrian artiopods. Philosophical Transactions of The Royal Society B Biological Sciences, 377, 1847. https://doi.org/10.1098/rstb.2021.0030.
27. Zhuang Y., Ran H., Li X., Feng C., Liu, Y.*. 2022. A new species of the iron maiden ant based on an alate female from mid-Cretaceous Burmese amber (Hymenoptera: Formicidae: †Zigrasimecia), Cretaceous Research, 130, 105056.https://doi.org/10.1016/j.cretres.2021.105056.
28. Liu, Y., Edgecombe, G.D.*, Schmidt, M., Bond, A.D., Melzer, R.R., Zhai, D., Mai, H., Zhang, M., Hou, X.*. 2021. Exites in Cambrian arthropods and homology of arthropod limb branches. Nature Communications, 12, 4619.https://doi.org/10.1038/s41467-021-24918-8.
29. Yang, X.*, Kimmig, J.*, Zhai, D., Liu, Y., Kimmig S.R., Peng, S. 2021. A juvenile-rich palaeocommunity of the lower Cambrian Chengjiang biota sheds light on palaeo-boom or palaeo-bust environments. Nature Ecology and Evolution, 5, 1082–1090. https://doi.org/10.1038/s41559-021-01490-4.
30. Schmidt, M.*, Liu, Y.*, Hou, X., Haug, J.T., Haug, C., Mai, H., and Melzer, R.R. 2021. Intraspecific variation in the Cambrian: new observations on the morphology of the Chengjiang euarthropod Sinoburius lunaris. BMC Ecology and Evolution, 21, 127. https://doi.org/10.1186/s12862-021-01854-1.
31. Xing, L., Liu, Y.*, McKellar, R. C.*, Luque, J., Li, G., Wang, Y., Yi, Q., Sun, R., Wang, E., Audo, D.*. 2021. The first shrimp preserved in mid-Cretaceous Kachin amber: systematics, palaeoecology, and taphonomy. Science Bulletin, 66 (17), 1723–1726.https://doi.org/10.1016/j.scib.2021.05.008.
32. Jin, C., Mai, H., Chen, H., Liu, Y., Hou, X., Wen, R. and Zhai, D. 2021. A new species of the Cambrian bivalved euarthropod Pectocaris with axially differentiated enditic armatures. Papers in Palaeontology, 7(4), 1781–1792. https://doi.org/10.1002/spp2.1362.
33. Schmidt, M.*, Liu, Y.*, Zhai, D., Hou, X., and Melzer, R. R. 2020. Moving legs: a workflow on how to generate a flexible endopod of the 518 million-year-old Chengjiang arthropod Ercaicunia multinodosa using 3d‐kinematics (Cambrian, China). Microscopy Research and Technique, 84(4), 695–704. https://doi.org/10.1002/jemt.23628.
34. O'Flynn, R.J., Audo, D., Williams, M., Zhai, D., and Liu, Y.* .2020. A new euarthropod with 'great appendage'-like frontal head limbs from the Chengjiang Lagersttäte, southwest China. Palaeontologia Electronica, 23(2), a36. https://doi.org/10.26879/1069.
35. Peng, S.*, Yang, X., Liu, Y., Zhu, X., Zhang, Y. 2020. Fulu biota, a new exceptionally-preserved Cambrian fossil assemblage from the Longha formation in southeastern Yunnan. Palaeoworld, 29(3), 453–461.https://doi.org/10.1016/j.palwor.2020.02.001.
36. Liu, Y.*, Ortega-Hernández, J.*, Zhai, D., Hou, X. 2020. A Reduced Labrum in a Cambrian Great-Appendage Euarthropod. Current Biology, 30, 3057–3061.https://doi.org/10.1016/j.cub.2020.05.085.
37. Liu, Y.*, Ortega-Hernández, J.*, Chen, H., Mai, H., Zhai, D., Hou, X. 2020. Computed tomography sheds new light on the affinities of the enigmatic euarthropod Jianshania furcatus from the early Cambrian Chengjiang biota. BMC Evolutionary Biology, 20, 62. https://doi.org/10.1186/s12862-020-01625-4.
38. Chen, H., Legg, D., Liu, Y., Hou, X.G. 2020. New data on the anatomy of fuxianhuiid arthropod Guangweicaris spinatus from the lower Cambrian Guanshan Biota, Yunnan, China. Acta Palaeontologica Polonica, 65 (1): 139–148.https://doi.org/10.4202/app.00508.2018.
39. Liu, Y.*, Lerosey-Aubril, R., Audo, D., Zhai, D., Mai, H., Ortega-Hernández. J.*. 2020. Occurrence of the eudemersal radiodont Cambroraster in the early Cambrian Chengjiang Lagerstätte and the diversity of hurdiid ecomorphotypes. Geological Magazine, 157 (7): 1200–1206. https://doi.org/10.1017/S0016756820000187.
40. Zhai, D., Edgecombe, G., Bond, A., Mai, H., Hou, X.*, Liu, Y.*. 2019. Fine-scale appendage structure of the Cambrian trilobitomorph Naraoia spinosa and its ontogenetic and ecological implications. Proceedings of the Royal Society B: Biological Sciences, 286, 20192371. https://doi.org/10.1098/rspb.2019.2371.
41. Zhai, D., Williams, M., Siveter, David J., Harvey, T.H.P., Sansom, R.S., Gabbott, S.E., Siveter, Derek J., Ma, X., Zhou, R., Liu, Y.*, Hou, X.*. 2019. Variation in appendages in early Cambrian bradoriids reveals a wide range of body plans in stem-euarthropods. Communications Biology, 2, 329. https://doi.org/10.1038/s42003-019-573-5.
42. Chen, X., Ortega-Hernández, J.*, Wolfe, J.M., Zhai, D., Hou, X., Chen, A., Mai, H., Liu, Y.*. 2019. The appendicular morphology of Sinoburius lunaris and the evolution of the artiopodan clade Xandarellida (Euarthropoda, early Cambrian) from South China. BMC Evolutionary Biology, 19, 165. https://doi.org/10.1186/s12862-019-1491-3.
43. Zhai, D., Ortega-Hernandez, J., Wolfe, J.M., Hou, X., Cao, C., Liu, Y.*. 2019. Three-Dimensionally Preserved Appendages in an Early Cambrian Stem-Group Pancrustacean. Current Biology, 29(1), 171–177.https://doi.org/10.1016/j.cub.2018.11.060.
44. Chen, A., Chen, H., Legg, D.A., Liu, Y.*, Hou, X. 2018. A redescription of Liangwangshania biloba Chen, 2005, from the Chengjiang biota (Cambrian, China), with a discussion of possible sexual dimorphism in fuxianhuiid arthropods. Arthropod Structure & Development, 47 (5), 552–561. https://doi.org/10.1016/j.asd.2018.08.001.
45. Hou, X., Williams, M., Sansom, R., Siveter, Derek J., Siveter, David J., Gabbott, S., Harvey, T.H.P., Cong, P., Liu, Y.*. 2018. A new xandarellid euarthropod from the Cambrian Chengjiang biota, Yunnan Province, China. Geological Magazine, 156(8), 1375–1384. https://doi.org/10.1017/S0016756818000730.
46. 赵婷; 侯先光; 翟大有; 吴迪; 陈红; 臧少刚; 刘煜, 2017. 显微CT技术在澄江生物群节肢动物研究中的应用——以Misszhouia longicaudata为例.古生物学报, 56 (04), 476–482.
47. 吴迪; 刘煜; 赵婷; 陈红; 侯先光*, 2017. 关山生物群中始海百合化石的再研究.古生物学报, 56 (04), 504–515.
48. Boyan, G., Liu, Y., Khalsa, S.K., Hartenstein, V. 2017. A conserved plan for wiring up the fan-shaped body in the grasshopper and Drosophila. Development Genes & Evolution, 227 (4), 253–269. https://doi.org/10.1007/s00427-017-0587-2.
49. He, Y., Cong, P., Liu, Y., Edgecombe, G.D., Hou, X.*. 2017. Telson morphology of Leanchoiliidae (Arthropoda: Megacheira) highlighted by a new Leanchoilia from the Cambrian Chengjiang biota. Alcheringa: An Australasian Journal of Palaeontology, 41 (4), 581–589.https://doi.org/10.1080/03115518.2017.1320425.
50. Liu, Y., Melzer, R.R., Haug, J.T., Haug, C., Briggs, D.E.G., Hörnig, M.K., He, Y., Hou, X.*. 2016. Three-dimensionally preserved minute larva of a great-appendage arthropod from the early Cambrian Chengjiang biota. PNAS, 113, 5542–5546.https://doi.org/10.1073/pnas.1522899113.
51. Yang, J., Ortega-Hernández, J., Butterfield, N.J., Liu, Y., Boyan, G.S., Hou, J., Lan, T., Zhang, X.*.2016. Fuxianhuiid ventral nerve cord and early nervous system evolution in Panarthropoda. PNAS, 113, 2988–2993.https://www.pnas.org/doi/10.1073/pnas.1522434113.
52. Boyan, G.*, Liu, Y. 2016. Development of the neurochemical architecture of the central complex. Frontiers in Behavioral Neuroscience, 10, 167.https://doi.org/10.3389/fnbeh.2016.00167.
53. Ehrhardt, E., Graf, P., Kleele, T., Liu, Y., Boyan, G.*. 2016. Fates of identified pioneer cells in the antennal nervous system of the grasshopper Schistocera gregaria. Arthropod, Structure & Development, 45, 23–30. https://doi.org/10.1016/j.asd.2015.11.001.
54. Strausfeld, N.J., Ma, X.*, Edgecombe, G.D., Fortey, R.A., Land, M.F., Liu, Y., Cong, P., Hou, X.*. 2016. Arthropod eyes: The early Cambrian fossil record and divergent evolution of visual systems. Arthropod, Structure & Development, 45, 152–172.https://doi.org/10.1016/j.asd.2015.07.005.
55. Liu,Y., Scholtz, G., Hou, X.*. 2015. When a 520 million-year-old Chengjiang fossil meets a modern micro-CT – a case study. Scientific Reports, 5, 12802.https://doi.org/10.1038/srep12802.
56. Boyan, G.*, Williams, L., Liu, Y. 2015. Conserved patterns of axogenesis in the protocerebral midline of panarthropod. Arthropod, Structure & Development, 44, 101–112. https://doi.org/10.1016/j.asd.2014.11.003.
57. Ehrhardt, E., Liu, Y., Boyan, G. 2015. Axogenesis in the antennal nervous system of the grasshopper Schistocera gregaria revisited: the base pioneers. Development, Genes & Evolution, 225, 39–45. https://doi.org/10.1007/s00427-014-0485-9.
58. Liu, Y., Haug, J.T., Haug, C., Briggs, D.E.G., Hou, X.*. 2014. A 520 million-year-old chelicerate larva. Nature Communications, 5: 4440.https://doi.org/10.1038/ncomms5440.
59. Boyan, G.*, Liu, Y. 2014a. Timelines in the insect brain: fates of identified neural stem cells generating the central complex in the grasshopper Schistocerca gregaria. Development, Genes & Evolution, 224, 37–51.https://doi.org/10.1007/s00427-013-0462-8.
60. Boyan, G.*, Liu, Y. 2014b. Dye coupling and immunostaining of astrocyte-like glia following intracellular injection of fluorochromes in brain slices of the grasshopper Schistocerca gregaria. Methods in Molecular Biology, 1082, 99.https://doi.org/10.1007/978-1-62703-655-9_7.
61. Liu,Y.*. 2013. Gliogenesis in the embryonic brain of the grasshopper Schistocerca gregaria with particular focus on the protocerebrum prior to mid-embryogenesis. Cell & Tissue Research, 354, 697–705. https://doi.org/10.1007/s00441-013-1682-y.
62.Liu,Y.* , Boyan, G. 2013. Glia associated with central complex lineages in the embryonic brain of the grasshopper Schistocerca gregaria. Development, Genes & Evolution, 223, 213–223. https://doi.org/10.1007/s00427-013-0439-7.
63. Boyan, G., Liu, Y., Loser, M. 2012. A cellular network of dye-coupled glia associated with the embryonic central complex in the grasshopper Schistocerca gregaria. Development, Genes & Evolution, 222, 125–138.https://doi.org/10.1007/s00427-012-0394-8.
64. Haug, J.T.*, Waloszek, D., Maas, A., Liu, Y., Haug, C. 2012. Functional morphology, ontogeny and evolution of mantis shrimp-like predators in the Cambrian. Palaeontology, 55, 369–399. https://doi.org/10.1111/j.1475-4983.2011.01124.x.
65. Boyan, G. *, Loser, M., Williams, L., Liu,Y. 2011. Astrocyte-like glia associated with the embryonic development of the central complex in the grasshopper Schistocerca gregaria. Development, Genes & Evolution, 221, 141–155.https://doi.org/10.1007/s00427-011-0366-4.
66.Liu,Y.* , Maas, A., Waloszek, D. 2010. Early embryonic development of the head region of Gryllus assimilis Fabricius, 1775 (Orthoptera, Insecta). Arthropod, Structure & Development, 39, 382–395. https://doi.org/10.1016/j.asd.2010.05.008.
67.Liu,Y.*, Maas, A., Waloszek, D. 2009. Early development of the anterior body region of the grey widow spider Latrodectus geometricus Koch, 1841 (Theridiidae, Araneae). Arthropod, Structure & Development, 38, 401–416.https://doi.org/10.1016/j.asd.2009.04.001.
68. Liu,Y., Bergström, J., Hou, X.*.2007. Chengjiang arthropod Leanchoilia illecebrosa(Hou, 1987) reconsidered. GFF, 129, 263–272. https://doi.org/10.1080/11035890701293263.